Who said "Gratitude is the healthiest of all human emotions."?

This quote is attributed to Herbert Boyer.

Gratitude is the healthiest of all human emotions. - Herbert Boyer

"Gratitude is the healthiest of all human emotions."

Herbert Boyer Gratitude

Molecular Biologist American Born 1936 (age 90)

"The future belongs to those who believe in the beauty of their dreams."

"Science is the most reliable way to understand the natural world."

"Life is what happens when you're busy making other plans."

"The only true wisdom is in knowing you know nothing."

"Gratitude makes life richer."

"A grateful heart is a happy heart."

"Respect those who came before us."

"Life's greatest treasures are the people we love"