Who said "The organism fulfills the program written in its genes."?

This quote is attributed to Wilhelm Johannsen.

The organism fulfills the program written in its genes. - Wilhelm Johannsen

"The organism fulfills the program written in its genes."

Wilhelm Johannsen Science

Geneticist and Botanist Danish 1857–1927

"Genetics is the science of heredity, and heredity is the fundamental principle of life."

"The gene is the unit of heredity, but it is not the unit of evolution."

"Nature does nothing suddenly, and gradualism is the law of change."

"Pure lines are the foundation of scientific breeding."

"Science helps us understand the world."

"I've witnessed the birth of science, and it continues to amaze me."

"I'm something of a scientist myself."

"The fusion process is self-sustaining now!"