Who said "Selection can only work with what heredity provides."?

This quote is attributed to Wilhelm Johannsen.

Selection can only work with what heredity provides. - Wilhelm Johannsen

"Selection can only work with what heredity provides."

Wilhelm Johannsen Nature

Geneticist and Botanist Danish 1857–1927

"Genetics is the science of heredity, and heredity is the fundamental principle of life."

"The gene is the unit of heredity, but it is not the unit of evolution."

"Nature does nothing suddenly, and gradualism is the law of change."

"Pure lines are the foundation of scientific breeding."

"Nature teaches us about beauty."

"Nature teaches us patience and wisdom"

"Sometimes it snows in April."

"Every town has a soul, and Twin Peaks has a troubled one."