Who said "Heredity and environment are two halves of the whole."?

This quote is attributed to Wilhelm Johannsen.

Heredity and environment are two halves of the whole. - Wilhelm Johannsen

"Heredity and environment are two halves of the whole."

Wilhelm Johannsen

Geneticist and Botanist Danish 1857–1927

"Genetics is the science of heredity, and heredity is the fundamental principle of life."

"The gene is the unit of heredity, but it is not the unit of evolution."

"Nature does nothing suddenly, and gradualism is the law of change."

"Pure lines are the foundation of scientific breeding."